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The 386 processor is a 32-bit processor, so it works in 4-byte chunks. This network shifts bytes around so the chunk is aligned correctly. Bits go down the vertical wires, across horizontal wires, and then out the bottom. Energizing different wires provides the desired shift and lines up the bytes.
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If this isn't enough about the 386 processor's prefetch unit, see my latest blog post. It has more photos, schematic diagrams, and lots of explanation: www.righto.com/2025/05/386-...