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In 1928, B.H. Danser had only flat 1894 herbarium sheets of Nepenthes clipeata from Borneo’s Gunung Kelam, yet deduced its cliff-wall habit from geometry: with a short, stiff tendril emerging mid-underside of a round shield leaf, a 30 cm pitcher had no clearance unless the leaf stood vertically.

When Mansur et al. (2021) roped onto Gunung Kelam’s granite cliffs after fires and poaching, they found just 18 wild plants—and among 14 in flower, 13 were male and 1 female. Their leaves stay N-limited (N:P 13.6) yet carry a distinct K-rich granite ionome (3.37% K) and dry-rim water economy.

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